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Bird-watching hotspots for flamingos include Djerba island off Tunisia's south-eastern coast, and Korba Lagoon further to the north.

Along with his wife and daughter, he is survived by another daughter, Djerba Goldfinger, and a grandchild.

In the attack on Djerba, a Tunisian naval guard shot and killed a colleague and two civilians - one of them French - as he tried to reach the 2,500-year-old Ghriba temple, one of Africa’s oldest synagogues.

Last Tuesday a National Guardsman killed two Jewish visitors and two policemen at the synagogue on Djerba island - Africa's oldest - before being shot dead.

The pilgrimage to the Djerba festival regularly draws hundreds of Jews from Europe and Israel and has operated under tight security since al Qaeda militants attacked the synagogue in 2002 with a truck bomb, killing 21 Western tourists.