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There was no training on the Geneva Convention, to which Russia and Ukraine are both signatories, and which prohibits the killing of people who have surrendered or no longer pose a threat.

"When people are still missing, how can they possibly complete the DNA process by tomorrow? What if my niece's remains have not even been found? The wait is killing us," he said.

Before the killing, President Emmanuel Macron had angered the right by saying they were too obsessed with crime, and not sufficiently interested in other issues like the environment.

Israel may calculate that the attacks and killings could unsettle the regime and open the way for a popular uprising.

Israel struck dozens of targets, including nuclear facilities, military sites and private residences, across Iran on Friday, killing a number of senior military commanders in what it called "Operation Rising Lion".