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A papery remnant of nucellus lines the inner face of the woody shell, and, as in cycadean seeds, the apical portion is readily separated as a cap covering the summit of the endosperm.

The remainder of the nucellus and the integuments of the ovules form the seed-coats.

There are thus two integuments to the nucellus, an outer and an inner.

The nucellus is a cellular tissue enveloping one large cell, the embryo-sac or macrospore.

Each ovule is enclosed at the base by an envelope or collar homologous with the lamina of a leaf; the fleshy and hard coats of the nucellus constitute a single integument.